ERA

Download the full-sized PDF of Gleichenia appianense sp. nov. (Gleicheniaceae): A permineralized rhizome and associated vegetative remains from the Eocene of Vancouver Island, British ColumbiaDownload the full-sized PDF

Analytics

Share

Permanent link (DOI): https://doi.org/10.7939/R3CR5NG78

Download

Export to: EndNote  |  Zotero  |  Mendeley

Communities

This file is in the following communities:

Biological Sciences, Department of

Collections

This file is in the following collections:

Journal Articles (Biological Sciences)

Gleichenia appianense sp. nov. (Gleicheniaceae): A permineralized rhizome and associated vegetative remains from the Eocene of Vancouver Island, British Columbia Open Access

Descriptions

Author or creator
Mindell, R. A.
Stockey, R. A.
Rothwell, G. W.
Beard, G.
Additional contributors
Subject/Keyword
Hymenophyllaceae
Fern
Lygodium
Eocene
Gleicheniaceae
Schizaeaceae
Type of item
Journal Article (Published)
Language
English
Place
Time
Description
A permineralized gleicheniaceous rhizome with an attached stipe base has been identified from the Appian Way locality on Vancouver Island, British Columbia, Canada. The specimens are preserved in Middle Eocene marine calcareous concretions that are studied using the cellulose acetate peel technique. The marginally mesarch rhizome has a vitalized protostele, protoxylem elements with helical wall thickenings, and scalariform metaxylem tracheids that occur in clusters. A continuous band of phloem and pericycle surrounds the xylem. The inner cortex consists of a layer of small-diameter sclerenchyma fibers, and the outer cortex is composed of larger parenchyma cells. A nodal island of sclerenchyma is present between the leaf trace and protostele at the level of frond-trace divergence. Long metaxylem tracheids in the frond trace show septa in longitudinal section that likely represent tyloses. Roots diverge from all sides of the rhizome with traces that run obliquely through the cortex. Numerous associated but isolated frond segments are found in the AppianWay nodules that show a pinched and inrolled C-shaped vascular trace and pseudodichotomous bifurcation. The frond traces have numerous protoxylem strands, and the general anatomy is comparable to that of gleicheniaceous stipes. The rhizome is described as a new species of Gleichenia, Gleichenia appianensis Mindell, Stockey, Rothwell, et Beard sp. nov., and represents the first record of Gleicheniaceae in the Tertiary of North America.
Date created
2006
DOI
doi:10.7939/R3CR5NG78
License information
Rights
© 2006 University of Chicago Press. This version of this article is open access and can be downloaded and shared. The original author(s) and source must be cited.
Citation for previous publication
Mindell, R.A., R.A. Stockey, G.W. Rothwell, and G. Beard. (2006). Gleichenia appianense sp. nov. (Gleicheniaceae): A permineralized rhizome and associated vegetative remains from the Eocene of Vancouver Island, British Columbia. International Journal of Plant Sciences, 167(3), 639-647. DOI: 10.1086/500997.
Source
Link to related item

File Details

Date Uploaded
Date Modified
2014-05-01T01:56:45.085+00:00
Audit Status
Audits have not yet been run on this file.
Characterization
File format: pdf (Portable Document Format)
Mime type: application/pdf
File size: 1889454
Last modified: 2015:10:12 16:43:57-06:00
Filename: IJOPS_167_2006_639.pdf
Original checksum: 9114eb90ef22c83538292863312349ae
Well formed: false
Valid: false
Status message: Invalid object number in cross-reference stream offset=1889419
File title: <italic>Gleichenia appianensis</italic> sp. nov. (Gleicheniaceae): A Permineralized Rhizome and Associated Vegetative Remains from the Eocene of Vancouver Island, British Columbia
File language: en
Activity of users you follow
User Activity Date