Download the full-sized PDF of Systematics and palaeobiology of the crested hadrosaurine Saurolophus from Canada and MongoliaDownload the full-sized PDF



Permanent link (DOI):


Export to: EndNote  |  Zotero  |  Mendeley


This file is in the following communities:

Graduate Studies and Research, Faculty of


This file is in the following collections:

Theses and Dissertations

Systematics and palaeobiology of the crested hadrosaurine Saurolophus from Canada and Mongolia Open Access


Other title
Horseshoe Canyon Formation
Nemegt Formation
Moreno Formation
skin impressions
Type of item
Degree grantor
University of Alberta
Author or creator
Bell, Phil
Supervisor and department
Currie, Philip (Biological Science)
Examining committee member and department
Lemelin, Pierre (Division of Anatomy)
Brinkman, Don (Royal Tyrrell Museum)
Horner, John (Museum of the Rockies)
Gingras, Murray (Earth and Atmospheric Science)
Caldwell, Michael (Biological Science)
Department of Biological Sciences

Date accepted
Graduation date
Doctor of Philosophy
Degree level
Reappraisal of Saurolophus confirms Saurolophus osborni and Saurolophus angustirostris as distinct species. Synapomorphies of Saurolophus include a spike-like pseudonarial crest formed by the nasals, frontals, and prefrontals; tripartite frontals with anteroventral and posterodorsal extensions that buttress the underside of the nasals; posterodorsal process on the prefrontal that buttresses the posterolateral edge of the nasal crest; and the presence of at least one supraorbital element. Phylogenetic analysis supports the close relationship between Saurolophus, Prosaurolophus, and Kerberosaurus. Material from the Maastrichtian Moreno Formation, California, previously assigned to Saurolophus is poorly preserved and lacks important diagnostic elements. This taxon shares traits of both Saurolophus and Edmontosaurus; however, as no unambiguous synapomorphies could be identified, this material is conservatively reassigned to Hadrosaurinae indet. The integument of Saurolophus varies significantly along the length of the body in both species. For the first time in the Dinosauria, soft tissue anatomy can be used to differentiate between species of single genus. A new standardised scheme for scale descriptions is also proposed. A Saurolophus bonebed from Mongolia represents the only dinosaur bonebed that preserves skin impressions. Ontogenetic evidence from this site suggests Barsboldia sicinskii, the only other hadrosaurid from the Nemegt Formation, is a junior synonym of Saurolophus angustirostris. An Edmontosaurus bonebed from the Horseshoe Canyon Formation (Alberta) contains nine animals including at least one Albertosaurus. The assemblage was subject to exposure, trampling, and rampant scavenging over a relatively short period (<1 year?). Future directions for hadrosaurid research are discussed.
License granted by Phil Bell ( on 2011-08-29T05:11:43Z (GMT): Permission is hereby granted to the University of Alberta Libraries to reproduce single copies of this thesis and to lend or sell such copies for private, scholarly or scientific research purposes only. Where the thesis is converted to, or otherwise made available in digital form, the University of Alberta will advise potential users of the thesis of the above terms. The author reserves all other publication and other rights in association with the copyright in the thesis, and except as herein provided, neither the thesis nor any substantial portion thereof may be printed or otherwise reproduced in any material form whatsoever without the author's prior written permission.
Citation for previous publication

File Details

Date Uploaded
Date Modified
Audit Status
Audits have not yet been run on this file.
File format: pdf (Portable Document Format)
Mime type: application/pdf
File size: 157320384
Last modified: 2015:10:22 06:13:19-06:00
Filename: Bell_Phil_Fall 2011.pdf
Original checksum: aa0ef3bc7962157fdb70534365341a52
Well formed: true
Valid: true
Status message: File header gives version as 1.4, but catalog dictionary gives version as 1.3
Status message: Too many fonts to report; some fonts omitted. Total fonts = 1469
File title: Microsoft Word - BELL PHD THESIS_2011.doc
File author: Phil Bell
Page count: 326
Activity of users you follow
User Activity Date